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LONGLEAF PINE


Longleaf pine (Pinus palustris) forests of the southern United States are declining and are in need of special attention from the conservation community (Means and Grow 1985, Noss 1989, Kelly and Bechtold 1990). Pre-European settlement longleaf pine forest may have extended up to 60,000,000 acres (Boyer 1990, Simberloff 1993, Frost 1993); yet, current estimates of remaining longleaf pine forests are just about 3,000,000 acres (Dennington and Farrar 1983, Engstrom et al. 1996). Therefore, many efforts are underway to reestablish longleaf pine on upland site (Moser 1996, Landers et al. 1995, USDA 1993b). A question that arises is the balance between management of mature, single-aged forests and uneven-aged timber because the uneven-aged timber management produces a forest more similar to the pre-settlement forests (Engstrom et al. 1996). The original forest matrix is thought to contain longleaf pine stands that vary greatly in their structure (Schwartz 1907, Wahlenberg 1946). One of the reasons for these stands that have a variety of ages and diameter classes is numerous small-scale disturbances (such as lightening strikes and varying intensity of fires) (Platt et al. 1988, Engstrom et al. 1996). Before European settlement fires were thought to have occurred as frequently as every 1-10 years (Chapman 1932a,b, 1950a,b, Christensen 1981, 1988).

Thus, fire is a recognized way of managing longleaf forests (Landers et al. 1995). However, there are problems in the resulting interactions between microclimate and physiology and ecology of regeneration of these forests (Croker 1990, Boyer and White 1990). These problems derive from a variety of sources. The species produces seed only irregularly (Boyer and White 1990). Competition among the roots for water may be a factor reducing the presence of seedlings in mature forest stands (Chapman 1909, Brown 1958). Numerous studies show that mature pine stands have few juveniles and that there is a inverse relationship between juvenile growth and overstory density (Pessin 1938, 1939, Heyward 1939, Smith 1955, Boyer 1963, 1974). Furthermore, the heavily weighted longleaf pine seeds do not fall very far from the parent tree (Croker and Boyer 1975). Recently, it has been shown that shelter woodsą methods are an appropriate method for regeneration (Croker and Boyer 1975, Neel 1993, Farrar 1993, 1996, Rudolph and Conner 1996). Because different disturbance patterns favor particular tree sizes, they can cause gaps in different sizes (Platt and Bradburn 1993, Palik and Patterson 1996).

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REFERENCES

Boyer, W.D., and J. B. White. 1990. Natural regeneration of longleaf pine. pp. 94-113 In Proc. of the Symposium on the Management of Longleaf Pine. Farrar RM (ed.) General Technical Report SO-75. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Boyer, W.D. 1963. Development of longleaf pine seedlings under parent trees. Research Paper SO-4. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Boyer, W. D. 1974. Impact of prescribed fires on mortality of released and unreleased longleaf pine seedlings. Research Note SO-128. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Boyer, W. D. 1990. Growing-season burns for control of hardwoods in longleaf pine stands. Research Paper SO-256. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Brown, C. L. 1958. Studies in growth and development of longleaf pine seedlings. Cambridge: Unpublished Harvard Univ Ph.D. Dissertation.

Chapman, H. H. 1909. A method of studying growth and yield of longleaf pine applied in Tyler Co., Texas. Proc: Soc. Am. For. Convention. 4: 207-220.

Chapman, H.H. 1932a. Is the longleaf type a climax? Ecology 13:328-334.

Chapman, H. H. 1932b. Some further relations of fire to longleaf pine. J. For. 30:602-604.

Chapman, H. H. 1950a. Lightning in the longleaf. Am. For. 56(1):10-11, 34.

Chapman, H. H. 1950b. Longleaf yellow pine owes its existence to fire. Coastal Cattleman 16(1):10-13.

Christensen, N. L. 1981. Fire regimes in southeastern ecosystems. pp. 112-130 In H. A. Mooney, T. M. Bonnickson, N. L. Christensen, J. E. Lotan, and W. A. Reiners (eds.) Fire regimes and ecosystem properties. General Technical Report WO-26. U.S.Department of Agriculture., Forest Service, Washington, D.C.

Christensen, N. L. 1988. Vegetation of the Southeastern Coastal Plain. pp. 317-364 In M. G. Barbour, and W.D. Billings (eds.) North American Terrestrial Vegetation. Cambridge: Cambridge Univ Pr.

Croker, T.C., and W.D. Boyer. 1975. Regenerating longleaf pine naturally. Research Paper SO-105. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Croker T.C. 1990. Longleaf pine: myths and facts. pp. 1-10 In Proc. Of the Symposium on the Management of Longleaf Pine. Farrar RM (ed.) General Technical Report SO-75. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Dennington, R. W. and R. M. Farrar. 1983. Longleaf pine management. Forestry Report R-8-FR-3. U.S. Department of Agriculture, Forest Service, Southern Region, Atlanta.

Engstrom, R. T., L. A. Brennan., W. L. Neel., R. M. Farrar, S. T. Lindeman, W. K. Moser, and S. M. Hermann. 1996. Silvicultural practices and red-cockaded woodpecker management: a reply to Rudolph and Conner. Wildl. Soc. Bull. 24(2):334-338.

Farrar, R. M. 1993. Growth and yield in naturally-regenerated pine stands. pp. 311-335 In S. Hermann (ed.) Proc. Of the Tall Timbers Fire Ecology Conference, No. 18, The Longleaf Pine Ecosystem: Ecology, restoration and management. Tallahassee, Fl: Tall Timbers Research Station.

Farrar, R. M. 1996. Fundamentals of Uneven-aged Management in Southern Pine. W. K. Moser and L. A. Brennan (eds.) Tallahassee, FL: Tall Timbers Research Station Misc Pub 9.

Frost, C. C. 1993. Four centuries of changing landscape patterns in the Longleaf Pine ecosystem. pp. 17-43 In S. Hermann (ed.) Proc. Of the Tall Timbers Fire Ecology Conference, No. 18, The longleaf Pine Ecosystem; Ecology, Restoration and Management. Tallahassee, FL: Tall Timbers Research Station.

Heyward, F. 1939. The relation of fire to stand composition of longleaf pine forests. Ecology 20:287-304.

Kelly, J. F., and W.A. Bechtold. 1990. The longleaf pine resource. pp. 11-22 In R. M. Farrar (ed.)Proc. Of the Symposium on the Management of Longleaf Pine. R. M. Farrar, R. M. (ed.) General Technical Report SO-75. U.S.Department of Agriculture., Forest Service, Southern Forest Experiment Station, New Orleans.

Landers, J. L., D. H. Van Lear, and W. D. Boyer. 1995. The longleaf pine forests in the Southeast: requiem or renaissance? J For. 93(11):39-44.

Means, B., and G. Grow. 1985. The endangered longleaf pine community. ENFO Bulletin. Winter Park, Fl: Environmental Information Center.

Moser, W. K. 1996. Extension Workshops on Longleaf Pine Management in the Southeastern United States: The View form an NGO. In: R. Beck and J. Johnson (eds.) Proc. Of the 1996 Meeting of IUFRO Working Group 6.06.03 Forestry Extension. Freising, Germany. In press.

Neel, W. L. 1993. An ecological approach to longleaf forestry. pp. 311-335 In S. Hermann (ed.) Proc. Of the Tall Timbers Fire Ecology Conference, No. 18, The Longleaf Pine Ecosystem: Ecology, restoration and management. Tallahassee, FL: Tall Timbers Research Station.

Noss, R. F. 1989. Longleaf pine and wiregrass: Keystone components of an endangered ecosystem. Nat Areas J. 9:234-35.

Palik, B. J., and N. Pederson. 1996. Overstory mortality and canopy disturbances in longleaf pine ecosystems. Can. J. For. Res. 26:2035-2047.

Pessin, L. J. 1938. The effect of vegetation on the growth of longleaf pine seedlings. Ecol Monogr. 8:115-149.

Pessin, L. J. 1939. Density of stocking and character of ground cover as a factor in longleaf pine reproduction. J. For. 37:255-258.

Platt, W. J., and S. L. Rathbun. 1993. Dynamics of an old-growth longleaf pine population. pp 275-298 In S. Hermann (ed.) Proc. Of the Tall Timbers Fire Ecology Conference, No. 18, The Longleaf Pine Ecosystem: Ecology, restoration, and management. Tallahassee FL: Tall Timbers Research Station.

Platt, W. J., G. W. Evans, and S. L. Rathbun. 1988. The population dynamics of a long-lived conifer (Pinus palustris). Am. Nat. 131(4):491-525.

Rudolph, D. C., Conner, R. N. 1996. Red-cockaded woodpeckers in silvicultural practice: Is even-aged silviculture preferable to uneven-aged? Wildl. Soc. Bull. 24(2):330-333.

Schwartz, G. F. 1907. The longleaf pine in virgin forest. J. Wiley, New York.

Simberloff, D. S. 1993. Species-area and fragmentation effects on old-growth forests: prospects for longleaf pine communities. pp. 1-13 In S. Hermann (ed.) Proc. Of the Tall Timbers Fire Ecology Conference, No. 18, The Longleaf Pine Ecosystem: Ecology, restoration, and management. Tallahassee FL: Tall Timbers Research Station.

Smith, L. F. 1955. Development of longleaf pine seedlings near large trees. J. For. 53:289-290.

USDA (U.S. Department of Agriculture). 1993b. Draft environmental impact statement for the management ofthe red-cockaded woodpecker and its habitat on national forests in the southern region. U.S. Department of Agriculture, Forest Service, Southern Region, Atlanta.

Wahlenberg, W. G. 1946. Longleaf pine: its use, ecology, regeneration, protection, growth, and management. Charles Lathrop Pack Foundation, Washington, D. C.


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